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Ph.D. in Botany, University of North Carolina at Chapel Hill, (1974)

M.S. in Botany, University of Wisconsin-Milwaukee, (1968)

B.A. in Biology, Beloit College, (1965)

Expertise

Research Interests

Taxonomy, ecology, evolution and control of parasitic weeds, especially Striga in the African Sahel and Orobanche and Cuscuta in the Middle East.Systematics and ecology of quillworts (Isoetes, Pteridophyta) in the southern United States, western Asia (with particular emphasis on Turkey, Syria, and Lebanon), and southern Africa.Plants of the Bible and Qur'an.Ecology and systematics of the Hydroraceae.Floristics of Iraq, Jordan, Lebanon, and Syria.Ethnobotany, especially of edible North American plants.

Articles

Bray, R. D., Schafran, P. W. and Musselman, L. J. (2018). Interesting, Provocative, and Enigmatic: Morphological Observations on Southeastern Quillworts (Isoetes Isoetaceae, Lycopodiophyta). Castanea 83 (2) , pp. 263-269.

Bolin, J. F. ., Lupton, D. and Musselman, L. J. (2018). Hydnora arabica sp. nov. (Aristolochiaceae), a new species from the Arabian Peninsula and a key to Hydnora. Phytotaxa 338 (1) , pp. 099-108.

Schafran, P. and Musselman, L. J. (2016). Isoetes mississippiensis: A new quillwort from Mississippi, USA. PhytoKeys Isoetes mississippiensis: A new quillwort from Mississippi, USA (74) , pp. 97-106.

Keskin, M. and Musselman, L. J. (2014). Türkiye'nin Çimeğreltisi [Isoetes L. (Isoetaceae)] türleri. Bağbahçe Bilim Dergisi 1 (2) , pp. 1-­‐13.

Bolin, J. F. and Musselman, L. J. (2013). Epitypification and ecological notes for the Malagasy holoparasite Hydnora esculenta Jum. and H. Perrier (Hydnoraceae). Nordic Journal of Botany 31 (3) , pp. 286-290.

Bolin, J. F., Bray, R. and Musselman, L. J. (2011). A New Species of Diploid Quillwort (Isoetes, Isoetaceae, Lycophyta) from Lebanon. Novon. Missouri Botanical Garden 21 , pp. 295-298.

Alarid, K., Bray, R. and Musselman, L. J. (2011). Microspore wall morphogenesis of Isoetes piedmontana. International Journal of Plant Science 172 (7).

Bolin, J. F., Tennakoon, K. U., Maass, E. and Musselman, L. J. (2010). Host specific germination of the root holoparasite Hydnora triceps (Hydnoraceae). Botany 87 (12) , pp. 1250-1254.

Musselman, L. J. and Al Zein, M. S. (2010). Isoetes duriei in Lebanon. American Fern Journal 99 (4) , pp. 333-334.

Chak, W. H., Tennakoon, K. U. and Musselman, L. J. (2010). The first report about dodders, the angiosperm parasitic genus Cuscuta (Yuncker) in Brunei Darussalam: A mystifying occurrence. Folia Malaysiana 11 (1) , pp. 13-24.

Bolin, J. F., Maass, E. and Musselman, L. J. (2009). Pollination biology of Hydnora Africana Thunb. (Hyndoraceae) in Namibia: Brood-site mimicry with insect imprisonment. International Journal of Plant Science 170 , pp. 157-163.

Musselman, L. J. and Bolin, J. F. (2008). New infestation of branched broomrape Orobanche ramosa L. (Orobanaceae) on black medic (Medicago lupulina L.) (Fabaceae) in Virginia. Plant Disease 92 (2) , pp. 315.

Bolin, J. F., Bray, R. D., Meskin, M. and Musselman, L. J. (2008). The genus Isoetes L. (Isoetaceae, Lycophyta) in South-Western Asia. Turkish Journal of Botany 32 , pp. 447-457.

Tennakoon, K. U., Bolin, J. F., Musselman, L. J. and Maas, E. (2007). Structural attributes of the hypogeous holoparasite Hydnora triceps Drege & Meyer (Hydnoraceae). American Journal of Botany 94 (9) , pp. 1439-1449.

Al-Zein, M. and Musselman, L. J. (2006). A western Asian genus honoring a North American pioneer botanist. Castanea Occasional Papers 2 , pp. 200-205.

Musselman, L. J. (2006). The botanical activities of George Edward Post (1838-1909). Archives of Natural History 33 (2) , pp. 282-301.

Maass, E. and Musselman, L. J. (2004). Hydnora triceps (Hydnoraceae) - First record in Namibia and first description of fruits. Dinteria 29 , pp. 1-10.

Musselman, L. J. and Saoud, N. S. (2004). The types of George Edward Post in Beirut and Geneva. Turkish Journal of Botany 28 , pp. 155-160.

Books

Leopold, D. J. (2020). Wildflowers of the Adirondacks .

. (2013). Parasitic Orobanchaceae Parasitic mechanisms and control strategies. Springer-Verlad.

Musselman, L. J. and Wiggins, H. J . (2013). The Quick Guide to Wild Edible Plants Easy to Pick, Easy to Prepare. Johns Hopkins University Press.

Musselman, L. J. and Knepper, D. A. (2012). Plants of Chesapeake Bay—An Illustrated Guide. Johns Hopkins University Press.

Musselman, L. J. (2011). Dictionary of Bible Plants. Cambridge University Press.

Musselman, L. J. and Keillor, G. (2007). Figs and Dates, Laurel and Myyrh: Plants of the Bible and Quran. Timber Press.

Book Chapters

Mohamed, K. Ibrahim. and Musseman, L. John. (2019). Striga. Flora of North America North of Mexico. Edited by Flora of North America Volume 17 Magnoliophyta: Tetrachondraceae to Orobanchaceae. (pp. Pages 508-510) Oxford: Oxford University Press.

Nickrent, D. Lee. and Musselman, L. J. (2016). Parasitic Plants Pages 266-288 Chapter 17 in: Plant Pathology Concepts and Laboratory Exercises. (pp. 266-288) Boca Raton, Florida: CRC Press.

(2016). Plants of the Holy Bible and the Holy Qur’an Pflanzen der Heiligen Bücher Bibel und Koran (pp. 19-20) Bonn: Bundesamt für Naturschutz.

Mohamed, K. I., Bolin, J. F., Musselman, L. J. and Peterson, A. T. (2007). Genetic Diversity of Striga and Implications for Control and Modeling Future Distributions DistributionsIntegrating New Technologies for Striga Control: Towards Ending the Witch-Hunt (pp. 71-84) World Publishing Co.

Yoder, J. I. and Musselman, L. J. (2006). Striga: A Subterranean Parasitic Angiosperm (Witchweed) Encyclopedia of Plant and Crop Science New York: Taylor and Francis.

Musselman, L. J. and Mohamed, K. I. (2006). Striga: Flora of Somalia Royal Botanic Gardens: Kew (pp. 287-289).

Musselman, L. J. (2004). Cuscutaceae Flowering Plants of the Neotropics (pp. 187-188) New York: New York Botanical Garden.

Musselman, L. J. (2004). Hydnoraceae Flowering Plants of the Neotropics (pp. 187-188) New York Botanical Garden.

Presentations

Musselman, L. John. (November 27, 2019). Managing striga weed and other parasitic weeds Oral Presentation Best Practices for Improving Nutrition in Highland Areas Kigali, Rwanda.

Musselman, L. John. (November 21, 2019). Parasitic Plants Lecture Virginia Native Plant Society Williamsburg.

Schafran, P. W., Flanders, N. P., Dotterer, T., Musselman, I. A. and Musselman, L. J. (April 5, 2019). Demographic and spatial analysis of Blackwater Ecologic Preserve (Isle of Wight Co, VA) provides insight into the origin of longleaf pine (Pinus palustris) stands Oral Presentation Association of Southeastern Biologists Memphis, Tennessee.

Schafran, P. W., Bolin, J. F., Taylor, W. Carl., Musselman, E. and Musselman, L. John. (April 5, 2019). High-throughput flow cytometry and DNA sequencing identifies over 20 potential new species of Isoetes (Isoetaceae, Lycopodiophyta) in the southeastern US Oral Presentation Association of Southeastern Biologists Memphis, Tennessee.

Heyd, L. Z., Schafran, P. W. and Musselman, L. J. (April 5, 2019). Sporophyll anatomy differences among four Isoetes species Oral Presentation Association of Southeastern Biologists Memphis, Tennessee.

Musselman, L. J. (February 12, 2019). Holy Botany: Plants of the Bible Lecture Fort Worth, Texas.

Musselman, L. John. (November 14, 2018). Parasitic Plants in African Agriculture: A growing problem Oral Presentation ECHO International Agriculture Conference Fort Myers, Florida.

Musselman, L. John. (October 24, 2018). The Blackwater Ecologic Preserve Poster Longleaf Reflections Alexandria, Louisiana.

Musselman, L. J. (August 8, 2018). African Witchweeds and Their Relatives—Biology, Sanitation, and Control Lecture Nutrition in Dry Land Areas Arusha, Tanzania.

Musselman, L. and Musselman, L. John. (April 5, 2018). An Exploration of the Ecology, Parasitism, and Control of the Root Parasite Agalinis fasciculata (Orobanchaceae) on Pinus taeda and P. elliottii Poster Association of Southeastern Biologists Myrtle Beach, South Carolina.

Flanders, N. P ., Amos, I., Schafran, P. W., Waag, D., Musselman, L. John. and Kolgan, L. (April 5, 2018). Relationships between tree density and the growth and survival of longleaf pine (Pinus palustris) at Blackwater Ecological Preserve in Isle of Wight County, Virginia Oral Presentation Association of Southeastern Biologists Myrtle Beach, South Carolina.

Schafran, P. W., Zimmer, E., Musselman, W. Carl. and Musselman, L. J. (April 5, 2018). The Isoetes flaccida complex in the southeastern United States: unrecognized species diversity in southern Georgia and northern Florida Poster Association of Southeastern Biologists Myrtle Beach, South Carolina.

Musselman, L. J. and Wilson, A. (April 5, 2018). The native root hemiparasite Agalinis fasciculata damages commercial pine plantations Oral Presentation Association of Southeastern Biologists Myrtle Beach, South Carolina.

Schafran, P. W., Zimmer, E. ., Musselman, W. Carl. and Musselman, L. J. (April 5, 2018). Unexpected genetic diversity in polyploid Isoetes revealed with PacBio DNA sequencing Peter Schafran Oral Presentation Association of Southeastern Biologists Myrtle Beach, South Carolina.

Flanders, N. P., Walters, E. L ., Randled, C. P . and Musselman, L. J . (June 28, 2017). The role of generalist avian frugivores in determining the distribution of the mistletoe Phoradendron leucarpum Oral Presentation 14th World Congress on Parasitic Plants Asilomar, Monterey, California.

Schafran, P. W ., Dotterer, T., Flanders, N. P. and Musselman, L. J. (November 2, 2016). Restoration and maintenance of the Northernmost Longleaf Pine Community, Blackwater Ecological Preserve, Isle of Wight Co., VA Poster Longleaf Alliance Savannah, Georgia.

Musselman, L. John. (October 18, 2016). Native Plants for Cordials and Aperitifs Oral Presentation Native Plants for Cordials and Aperitifs Norfolk, Virginia .

Musselman, L. John. and Schafran, P. W. (October 14, 2016). Cordials and Aperitifs from Native Plants Oral Presentation Cordials and Aperitifs from Native Plants Hockkesin, Delaware.

Musselman, L. John. (August 23, 2016). Feel the Burn Oral Presentation Cruise, Alaska.

Musselman, L. John. (August 22, 2016). The Prophet's Pharmacy Oral Presentation Cruise, Alaska.

Musselman, L. John. (August 21, 2016). Into the Wild Oral Presentation Cruise, Alaska.

Musselman, L. John. (August 18, 2016). Plants Acting Like Animals Oral Presentation Alaska, cruise.

Musselma, L. John. (August 3, 2016). Root Parasites of Southern Forests Oral Presentation Rayonier Corporation Foresters Yule, Georgia.

Huisa, K., Schafran, P. W. and Musselman, L. J. (August 1, 2016). Analysis of morphological and anatomical characteristics of stomata of Isoetes hyemalis Oral Presentation Botanical Society of America Savannah, Georgia.

Schafran, P. W. and Musselman, L. J. (August 1, 2016). Molecular phylogeny of diploid Isoetes (Isoetaceae, Lycophyta) in the southeastern United States based on low-copy nuclear markers and whole chloroplast genomes. Oral Presentation Botanical Society of America Savannah, Georgia.

Musselman, L. John. (July 30, 2016). Edible Wild Plants of Virginia Lecture Edible Wild Plants Oswego, New York.

Musselman, L. John. (June 26, 2016). Plants that Act Like Animals Lecture Virginia Master Gardener College VPI, Blacksburg, VA.

Musselman, L. John. (June 25, 2016). Edible Wild Plants of Virginia Lecture Virginia Master Gardener College VPI, Blacksburg, VA.

Musselman, L. John. (May 5, 2016). The Tree that Built Tidewater: The neglected history of longleaf pine in Virginia. Lecture The Growing Interest in Longleaf Pine--longleaf pine workshop Paul D Camp Community College, Franklin, Virginia.

Musselman, L. john., Peter, S. W. and Rebecca, B. D. (April 2, 2016). Rhizomorphs, scales, and sporelings of Isoetes Oral Presentation Association of Southeastern Biologists Concord, NC.

Schafran, P. W., W , T. Carl., Rebecca, B. D., Jay, B. F. and Lytton, M. John. (April 2, 2016). Towards a phylogeny of Isoetes in the Southeastern United States Oral Presentation Association of Southeastern Biologists Concord, NC.

Flanders, N. F., Randle, C., Walters, E. and Musselman, L. John. (April 1, 2016). Role of generalist avian frugivores in determining the distribution of the mistletoe Phoradendron leucarpum Oral Presentation association of Southeastern Biologists concord, NC.

Musselman, L. John. (December 22, 2015). Holy Botany: Ethnobotany of the Holy Writings Lecture Atelier Tirana, Albania.

Musselman, L. John. (August 30, 2015). The Tree that Built Tidewater Oral Presentation Annual Landowner Retreat Wakefield, Virginia.

Musselman, L. John., Bolin, J. F., Al Rahbi, S. and Al-Hinai, A. (July 7, 2015). Taxonomy and Phylogenetics of Hydnoraceae: Recent Research Oral Presentation International Parasitic Plant Society 13th Congress Kunming, China.

Musselman, L. John. (May 27, 2015). Ethnobotany of Plants in the Holy Writings Oral Presentation Sultan Omar ‘Ali Saifuddien Centre for Islamic Studies, Universiti Brunei Darussalam.

Musselman, L. John. (April 3, 2015). A Review of the Taxonomy and Phylogenetics of the Hydnoraceae and a potentially new species from Oman Oral Presentation Association of Southeastern Biologists Chattanooga, Tennessee .

Musselman, L. John. (November 12, 2014). The Tree that Built Tidewater--The Forgotten Role of Longleaf Pine in Virginia History Keynote/Plenary Address Conservation Forum ������.

Musselman, L. John. (September 26, 2014). Longleaf pine Other Field Trip Blackwater Preserve.

Musselman, L. John. (May , 2014). Edible Wild Plants Other Edible Wild Plants Wilmington, Delaware.

Musselman, L. John. (May , 2014). Edible Wild Plants Lecture Oswego, New York.

Musselman, L. John. (April , 2014). Environmental Ethics in the Bible Lecture Islamic Perspectives on the Environment Doha, Qatar.

Matthews, R. A., Musselman, L. John., Taylor, W. Carl. and Schafran, P. W. (April 5, 2014). Isoetes snowii, a New Diploid Quillwort from the Southeastern United States Oral Presentation Association of Southeastern Biologists Spartanburg, South Carolina.

Schafran, P. W., Keenan, J., Jones, M., Bolin, J. F. and Musselman, L. John. (April 4, 2014). Germination and Development of Hazel Dodder Cuscuta corylii (Convolvulaceae) Oral Presentation Association of Southeastern Biologists Spartanburg, South Carolina.

Schafran, P. W., Keenan, J., Jones, M., Bolin, J. F. and Musselman, L. John. (April 4, 2014). Germination and Development of Hazel Dodder Cuscuta corylii (Convolvulaceae) Oral Presentation Association of Southeastern Biologists Spartanburg, South Carolina.

Schafran, P. W., Keenan, J., Jones, M. and Musselman, L. John. (April 4, 2014). In Vitro Seed Germination and Development of Calopogon pallidus (Orchidaceae) from Virginia Poster Association of Southeastern Biologists Spartanburg, South Carolina.

Musselman, L. John ., Bray, R., Schafran, P. W. and Taylor, W. Carl. (April 4, 2014). Misconceptions about Quillworts (Isoetes, Isoetaceae, Lycophyta) Oral Presentation Association of Southeastern Biologists Spartanburg, South Carolina.

Musselman, L. John ., Bray, R., Schafran, P. W. and Taylor, W. Carl. (April 4, 2014). Misconceptions about Quillworts (Isoetes, Isoetaceae, Lycophyta) Oral Presentation Association of Southeastern Biologists Spartanburg, South Carolina.

Schafran, P. W., Taylor, W. Carl., Bray, R. and Musselman, L. John. (April 4, 2014). Systematics of the Genus Isoetes (Isoetaceae, Lycophyta) in the Southeastern United States Oral Presentation Association of Southeastern Biologists Spartanburg, South Carolina.

Schafran, P. W., Taylor, W. Carl., Bray, R. and Musselman, L. John. (April 4, 2014). Systematics of the Genus Isoetes (Isoetaceae, Lycophyta) in the Southeastern United States Oral Presentation Association of Southeastern Biologists Spartanburg, South Carolina.

Musselman, L. John. (October 8, 2013). Holy Botany: Plants of the Bible Oral Presentation Brandeis University Waltham, Massachusets.

Musselman, L. John. (August , 2013). Four presentations Lecture A Prairie Home Companion cruise Cruise.

Musselman, L. John. (August , 2013). Three lectures: Deep Ecology, Why Mediterranean Herbs Burn, What Mustard Would Jesus Use? Lecture A Prairie Home Companion cruise Mediterranean .

Musselman, L. John. (July , 2013). The Role of Plants in Folklore and Religion Lecture SUNY College of Environmental Science and Forestry Cranberry Lake, New York.

Musselman, L. John. and Wiggins, H. J. (July 19, 2013). Quick Guide to Edible Plants Lecture Irvine Nature Center Irvine, Maryland.

Musselman, L. J. and Knepper, D. A . (July 13, 2013). Plants of Chesapeake Bay Lecture Adkins Arboretum Ridgely, Maryland.

Schafran, P. W., Wiggins, H. J . and Musselman, L. J. (April 6, 2012). The true identity of tuckahoe Oral Presentation Association of Southeastern Biologists University of Georgia, Athens.

Musselman, L. John. (October 11, 2011). The Strangest Plant in the World! Emirates Natural History Group Al Ein, United Arab Emirates.

Musselman, L. John. (October 5, 2011). Finding a Way Round--Plants that Eat Other Plants University of Brunei Faculty of Science, University of Brunei, Bandar Seri Begawan, Brunei Darussalam.

Musselman, L. J . (August 20, 2011). The Charm and Challenges of Bible Plant Research Silverton, OR.

Musselman, L. J. (August 3, 2011). The Strangest Plant in the World! Cranberry Lake, NY.

Musselman, L. John. (July , 2011). Three lectures on plants A Prairie Home Companion cruise Boston-Montreal.

Musselman, L. John., Maass, E. and Bolin, J. F. (June , 2011). Flora Biology of Hydnora abyssinica--New insights from southern Namibia Eleventh Congress on Parasitic Plants Martin Franca, Italy.

Musselman, L. John., Bolin, J. F. and Maass, E. (June , 2011). The Genus Hydnora in Southern Africa and Madagascar Eleventh World Congress on Parasitic Plants Martin Franca, Italy.

Musselman, L. John. (May , 2011). Parasitic Angiosperms College Seminar Sulaimani, Iraq.

Musselman, L. John. (April , 2011). Parasitic Angiosperms Spring Symposium Wilmore, KY.

Musselman, L. John. (August , 2010). Plants of the Bible Seminar Cranberry Lake Biological Station, New York.

Boley, J. F., Motley, T., Maass, E. and Musselman, L. J. ( 2009). Molecular phylogenetic relationships and a revised taxonomy of the holoparasitic family Hydnoraceae 9th International Parasitic Plants Symposium Izmir, Turkey.

Musselman, L. J. (October , 2009). Brunei Natural History Society Bandar Seri Begawan.

Musselman, L. J. (March , 2009). The Role of Plants in Religion and Folklore Botanical Society of Nigeria, Kano .

Musselman, L. J. (March , 2009). History of Botanical Exploration in the Middle East Flora of Iraq Sulimaniya, Iraq.

Musselman, L. J. (March , 2009). Botanical Exploration in the Middle East Flora of Iraq Organizational Meeting Twin Rivers Institute, Sulimaniya, Iraq.

Musselman, L. J. (March , 2009). Forest Management by Fire University of Sulimaniya, Sulimaniya, Iraq.

Musselman, L. J. (February , 2009). (Plenary Speaker) The Quranic Garden Quranic Garden, Doha, Qatar.

Bolin, J. F., Bray, R. D., Taylor, W. C. and Musselman, L. J. ( 2008). Unraveling the reticulate evolutionary history of the Isoetes hyemalis complex Botanical Society of America Vancouver, Canada.

Musselman, L. J. (October , 2008). Biodiversity in the Middle East (Plenary Speaker) Aqaba, Jordan.

Bolin, J. F., Tennakoon, K. U. and Musselman, L. J. ( 2007). Stable isotope and mineral relationships in the root holoparasite Hydnora (Hydnoraceae) 8th International Parasitic Plants Symposium Charlottesville, VA.

Bolin, J. F., Tennakoon, K. U. and Musselman, L. J. ( 2007). Stable isotope and mineral relationships in the root holoparasite Hydnora (Hydnoraceae) Association of Southeastern Biologists Columbia, SC.

Bolin, J. F., Tennakoon, K. U. and Musselman, L. J. ( 2007). Stable isotope and mineral relationships in the root holoparasite Hydnora (Hydnoraceae) Botanical Society of America Chicago, IL.

• 2019: Meritorious Teaching Award, Association of Southeastern Biologists
• 2014: Fulbright Specialist Program, Fulbright
• 2014: TEDx Speaker, TEDx
• 2004: Provost's Award for Leadership in International Education, ������
• 2001: Fellow, The International Parasitic Plant Society, Nantes, France
• 1999: Partnership Award, Virginia Chapter, The Nature Conservancy
• 1998: Induction into Phi Kappa Phi, Phi Kappa Phi
• 1997: Fullbright Awards,
• 1986: Faculty Research Award, ������
• 1986: Fullbright Awards,
• 1982: Fullbright Awards,
• 1972: Elected Fellow, Linnean Society of London